The first two stomachs of a ruminant comprised of the reticulum and the rumen. A microbial population lives in the rumen and enable the cow to digest dietary fiber. The digesta in the rumen and the reticulum is being exchanged once about every 50 to 60 seconds under the influence of a rhythmic cycle of contraction which also results in passage of some digesta from the reticulum into the omasum through the reticulo-omasal orifice.